Okay, they're not exactly the main villans, but they're still something that gets in your way.
I always think of Metroids as major antagonists. Sure, in Metroid II they were the "villan," but in all the other Metroid games, the Metroids were there to attempt to stop you from moving forward. They have a very creepy high-pitched squeal, are highly intelligent...and they fly around looking to latch onto your skull.
................Did I mention that they grow bigger if they suck energy from you?
Antagonists
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